{"data":{"id":"10.6084/m9.figshare.1252075.v1","type":"dois","attributes":{"doi":"10.6084/m9.figshare.1252075.v1","prefix":"10.6084","suffix":"m9.figshare.1252075.v1","identifiers":[],"alternateIdentifiers":[],"creators":[{"name":"Black, Karen H.","nameType":"Personal","givenName":"Karen H.","familyName":"Black","affiliation":[],"nameIdentifiers":[]},{"name":"Louys, Julien","nameType":"Personal","givenName":"Julien","familyName":"Louys","affiliation":[],"nameIdentifiers":[]},{"name":"Price, Gilbert J.","nameType":"Personal","givenName":"Gilbert J.","familyName":"Price","affiliation":[],"nameIdentifiers":[]}],"titles":[{"title":"Understanding morphological variation in the extant koala as a framework for identification of species boundaries in extinct koalas (Phascolarctidae; Marsupialia)"}],"publisher":"Taylor \u0026 Francis","container":{},"publicationYear":2014,"subjects":[{"subject":"110309 Infectious Diseases","subjectScheme":"FOR"},{"subject":"FOS: Health sciences","schemeUri":"http://www.oecd.org/science/inno/38235147.pdf","subjectScheme":"Fields of Science and Technology (FOS)"},{"subject":"FOS: Health sciences","subjectScheme":"Fields of Science and Technology (FOS)"},{"subject":"Biological Sciences"},{"subject":"Ecology"},{"subject":"FOS: Biological sciences","schemeUri":"http://www.oecd.org/science/inno/38235147.pdf","subjectScheme":"Fields of Science and Technology (FOS)"},{"subject":"FOS: Biological sciences","subjectScheme":"Fields of Science and Technology (FOS)"},{"subject":"Earth and Environmental Sciences"},{"subject":"Evolutionary Biology"},{"subject":"Molecular Biology"},{"subject":"Genetics"}],"contributors":[],"dates":[{"date":"2014-02-17","dateType":"Created"},{"date":"2016-01-19","dateType":"Updated"},{"date":"2014","dateType":"Issued"}],"language":null,"types":{"ris":"DATA","bibtex":"misc","citeproc":"dataset","schemaOrg":"Dataset","resourceType":"Dataset","resourceTypeGeneral":"Dataset"},"relatedIdentifiers":[{"relationType":"IsSupplementTo","relatedIdentifier":"10.1080/14772019.2013.768304","relatedIdentifierType":"DOI"},{"relationType":"IsIdenticalTo","relatedIdentifier":"10.6084/m9.figshare.1252075","relatedIdentifierType":"DOI"}],"relatedItems":[],"sizes":["52358 Bytes"],"formats":[],"version":null,"rightsList":[{"rights":"Creative Commons Attribution 4.0 International","rightsUri":"https://creativecommons.org/licenses/by/4.0/legalcode","schemeUri":"https://spdx.org/licenses/","rightsIdentifier":"cc-by-4.0","rightsIdentifierScheme":"SPDX"}],"descriptions":[{"description":"We document morphological variation (both geographical and sexual) in the dentition of the extant koala, \u003ci\u003ePhascolarctos cinereus\u003c/i\u003e, in order to facilitate discrimination of species boundaries in extinct phascolarctids. Considerable variation is evident in dental structures previously used to diagnose several phascolarctid fossil species. Consistent patterns of morphological variation are not evident between sexes or geographic regions, with variation as great between samples as within them. Metric variation is evident between the sexes in upper molar dimensions with Victorian (southern) males significantly larger than Victorian females, although this is not reflected in lower molar dimensions or in the Queensland (northern) sample. Male koalas from southern populations generally display significantly larger molars than their northern counterparts; however this trend is not evident in female upper molar dimensions. In both males and females, some, but not all, lower molar dimensions are larger in southern populations than northern. In light of these results, a systematic revision of species of \u003ci\u003eLitokoala\u003c/i\u003e suggests \u003ci\u003eL.\u003c/i\u003e ‘\u003ci\u003edicktedfordi\u003c/i\u003e’ is a junior synonym of \u003ci\u003eL. kutjamarpensis\u003c/i\u003e, and the poorly known \u003ci\u003eL. thurmerae\u003c/i\u003e is regarded to be a \u003ci\u003enomen dubium\u003c/i\u003e. Further, we describe a partial cranium of a new species of koala from Early Miocene sediments in the Riversleigh World Heritage Area, northern Australia. \u003ci\u003eLitokoala dicksmithi\u003c/i\u003e sp. nov. is the fifth koala species recorded from the diverse rainforest assemblages of Riversleigh and the third species referred to the Oligo-Miocene genus \u003ci\u003eLitokoala\u003c/i\u003e. Aspects of cranial morphology, including a shortened robust rostrum and broad, irregular nasal aperture, confirm placement of \u003ci\u003eLitokoala\u003c/i\u003e as sister taxon to the modern genus \u003ci\u003ePhascolarctos\u003c/i\u003e. Relatively large orbits and small body size suggest the possibility that \u003ci\u003eL. dicksmithi\u003c/i\u003e was nocturnal, had enhanced visual acuity, and was a more agile arboreal species than the relatively sedentary extant koala.http://zoobank.org/urn:lsid:zoobank.org:pub:EE5D13C1-47BB-4432-B281-16398A1781E1","descriptionType":"Abstract"}],"geoLocations":[],"fundingReferences":[],"xml":"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","url":"https://tandf.figshare.com/articles/dataset/Understanding_morphological_variation_in_the_extant_koala_as_a_framework_for_identification_of_species_boundaries_in_extinct_koalas_Phascolarctidae_Marsupialia_/1252075/1","contentUrl":null,"metadataVersion":2,"schemaVersion":"http://datacite.org/schema/kernel-4","source":"mds","isActive":true,"state":"findable","reason":null,"viewCount":0,"viewsOverTime":[],"downloadCount":0,"downloadsOverTime":[],"referenceCount":0,"citationCount":0,"citationsOverTime":[],"partCount":0,"partOfCount":0,"versionCount":0,"versionOfCount":0,"created":"2016-01-19T11:22:33.000Z","registered":"2016-01-19T11:22:34.000Z","published":"2014","updated":"2020-09-04T20:19:46.000Z"},"relationships":{"client":{"data":{"id":"figshare.ars","type":"clients"}},"provider":{"data":{"id":"otjm","type":"providers"}},"media":{"data":{"id":"10.6084/m9.figshare.1252075.v1","type":"media"}},"references":{"data":[]},"citations":{"data":[]},"parts":{"data":[]},"partOf":{"data":[]},"versions":{"data":[]},"versionOf":{"data":[]}}}}