{"data":{"id":"10.6084/m9.figshare.1165481","type":"dois","attributes":{"doi":"10.6084/m9.figshare.1165481","prefix":"10.6084","suffix":"m9.figshare.1165481","identifiers":[],"alternateIdentifiers":[],"creators":[{"name":"Gorscak, Eric","nameType":"Personal","givenName":"Eric","familyName":"Gorscak","affiliation":[],"nameIdentifiers":[]},{"name":"O'Connor, Patrick M.","nameType":"Personal","givenName":"Patrick M.","familyName":"O'Connor","affiliation":[],"nameIdentifiers":[]},{"name":"Stevens, Nancy J.","nameType":"Personal","givenName":"Nancy J.","familyName":"Stevens","affiliation":[],"nameIdentifiers":[]},{"name":"Roberts, Eric M.","nameType":"Personal","givenName":"Eric M.","familyName":"Roberts","affiliation":[],"nameIdentifiers":[]}],"titles":[{"title":"The basal titanosaurian \u003ci\u003eRukwatitan bisepultus\u003c/i\u003e (Dinosauria, Sauropoda) from the middle Cretaceous Galula Formation, Rukwa Rift Basin, southwestern Tanzania"}],"publisher":"Taylor \u0026 Francis","container":{},"publicationYear":2014,"subjects":[{"subject":"Cancer"},{"subject":"Biological Sciences"},{"subject":"Ecology"},{"subject":"FOS: Biological sciences","schemeUri":"http://www.oecd.org/science/inno/38235147.pdf","subjectScheme":"Fields of Science and Technology (FOS)"},{"subject":"FOS: Biological sciences","subjectScheme":"Fields of Science and Technology (FOS)"},{"subject":"Earth and Environmental Sciences"},{"subject":"Evolutionary Biology"},{"subject":"Physiology"}],"contributors":[],"dates":[{"date":"2014-09-09","dateType":"Created"},{"date":"2015-12-02","dateType":"Updated"},{"date":"2014","dateType":"Issued"}],"language":null,"types":{"ris":"DATA","bibtex":"misc","citeproc":"dataset","schemaOrg":"Dataset","resourceType":"Dataset","resourceTypeGeneral":"Dataset"},"relatedIdentifiers":[{"relationType":"IsSupplementTo","relatedIdentifier":"10.1080/02724634.2014.845568","relatedIdentifierType":"DOI"}],"relatedItems":[],"sizes":["148631 Bytes"],"formats":[],"version":null,"rightsList":[{"rights":"Creative Commons Attribution 4.0 International","rightsUri":"https://creativecommons.org/licenses/by/4.0/legalcode","schemeUri":"https://spdx.org/licenses/","rightsIdentifier":"cc-by-4.0","rightsIdentifierScheme":"SPDX"}],"descriptions":[{"description":"ABSTRACTWhereas titanosaurians represent the most diverse and cosmopolitan clade of Cretaceous sauropod dinosaurs, they remain rare components of Cretaceous African faunas. Currently recognized continental African titanosaurians include \u003ci\u003eAegyptosaurus baharijensis\u003c/i\u003e and \u003ci\u003eParalititan stromeri\u003c/i\u003e from early Upper Cretaceous deposits near Bahariya Oasis, Egypt, and \u003ci\u003eMalawisaurus dixeyi\u003c/i\u003e and \u003ci\u003eKarongasaurus gittelmani\u003c/i\u003e from the Lower Cretaceous (∼Aptian) Dinosaur Beds of Malawi, in addition to several undesignated and fragmentary forms across the continent. Here, we describe a new titanosaurian taxon, \u003ci\u003eRukwatitan bisepultus\u003c/i\u003e, on the basis of a partial, semiarticulated postcranial skeleton recovered from the middle Cretaceous Galula Formation in southwestern Tanzania. Unique to \u003ci\u003eRukwatitan\u003c/i\u003e are carotid processes on posterior cervical vertebrae, a deep coracobrachialis fossa and subquadrangular cross-section of the humerus, and a slender, curved, teardrop-shaped pubic peduncle on the ilium. Parsimony and Bayesian phylogenetic analyses of 35 sauropod taxa congruently place \u003ci\u003eRukwatitan\u003c/i\u003e as a non-lithostrotian titanosaurian, a relationship supported by cervical vertebrae with undivided pleurocoels and strongly procoelous anterior caudal vertebrae. \u003ci\u003eRukwatitan\u003c/i\u003e differs from the potentially penecontemporaneous and geographically proximate \u003ci\u003eMalawisaurus\u003c/i\u003e by exhibiting weakly developed chevron articulations and posteriorly inclined neural spines on the middle caudal vertebrae, a proximally robust and distally unexpanded humerus, and an anteroventrally elongated coracoid. Similar to biogeographic patterns identified in certain crocodyliform clades (e.g., small-bodied notosuchians), titanosaurians on continental Africa appear to exhibit a regional (e.g., southern versus northern Africa), rather than a continental- or supercontinental-level signal.SUPPLEMENTAL DATA—Supplemental materials are available for this article for free at www.tandfonline.com/UJVP","descriptionType":"Abstract"}],"geoLocations":[],"fundingReferences":[],"xml":"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